A spinach plant having an allele designated alpha-WOLF 23 encoding a protein conferring resistance to downy mildew and Cucumber Mosaic Virus, wherein said allele has: a nucleotide sequence having a coding sequence having at least 97% sequence identity to SEQ ID NO: 10; a nucleotide sequence encoding a protein having an amino acid sequence having at least 93.5% sequence identity to SEQ ID NO: 11; a nucleotide sequence encoding an LRR domain, said nucleotide sequence having at least 94.5% sequence identity to SEQ ID NO: 12; or a nucleotide sequence encoding an LRR domain having an amino acid sequence having at least 93% sequence similarity to SEQ ID NO: 13; and the plant does not include a second alpha-WOLF allele having a nucleotide having a coding sequence according to SEQ ID NO: 14.
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